Battle of Backbone Mountain

Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.^[1][2][3]

Families and genera

After Prokop et al. (2014), Guan et al. (2016) and Nel & Poschmann (2021):^[1][4][5]

• †Anthracoptilidae Handlirsch, 1922 (Synonyms: Strephocladidae Martynov, 1938; Strephoneuridae Martynov, 1940; ?Adiphlebiidae Handlirsch, 1906)
  • ?†Adiphlebia Scudder, 1885
  • †Afrocladus Nel, Garrouste & Prokop, 2015^[6]
  • †Anthracoptilus Lameere, 1917
  • †Carrizocladus Rasnitsyn in Rasnitsyn et al., 2004
  • †Graticladus Novokshonov and Aristov, 2004
  • †Homocladus Carpenter, 1966
  • †Jarmilacladus Rasnitsyn and Aristov, 2004
  • †Lodevocladus Prokop et al., 2015^[7]
  • †Mesoptilus Lameere, 1917
  • †Mycteroptila Rasnitsyn, 1977
  • †Paracladus Carpenter, 1966
  • †Pruvostia Bolton, 1921
  • †Pseudomesoptilus Guan et al., 2016
  • †Rhinomaloptila Rasnitsyn, 1977
  • †Spargoptilon Kukalová, 1965
  • †Strephocladus Scudder, 1885
  • †Strephoneura Martynov, 1940
  • †Strephoptilus Rasnitsyn and Aristov, 2013
  • †Westphaloptilus Guan et al., 2016
• †Blattinopsidae Bolton, 1925 (Synonym: Klebsiellidae Handlirsch, 1906)
  • †Alienus Handlirsch, 1906
  • †Avionblattinopsis Quispe et al., 2021^[8]
  • †Balduriella Meunier, 1925
  • †Blattinopsis Giebel, 1867
  • †Glaphyrophlebia Handlirsch, 1906
  • †Klebsiella Meunier, 1908
  • †Microblattina Scudder, 1895
  • †Protoblattiniella Meunier, 1912
  • †Rhipidioptera Brongniart, 1894
  • †Stephanopsis Kukalová, 1958
  • †Westphaloblattinopsis Béthoux and Jarzembowski, 2010
• †Paoliidae Handlirsch, 1906 (Synonyms: Ideliidae Zalessky, 1928; Herbstialidae Schmidt, 1953; Permulidae Aristov & Storozhenko, 2011; Permotermopsidae Martynov, 1937)
  • †Acropermula Aristov and Storozhenko, 2011
  • †Aenigmidelia Sharov, 1961
  • †Archidelia Sharov, 1961
  • †Carbonidelia Nel, Garrouste & Jouault, 2023^[9]
  • †Darekia Prokop et al., 2012
  • †Herbstiala Schmidt, 1952
  • †Holasicia Kukalová, 1958
  • †Idelina Storozhenko, 1992
  • †Kemperala Brauckmann, 1984
  • †Khosaridelia Storozhenko, 1992
  • †Kortshakolia Sharov, 1961
  • †Kochopteron Brauckmann, 1984
  • †Mertovia Prokop and Nel, 2007
  • †Micaidelia Aristov 2004
  • †Mongoloidelia Storozhenko, 1992
  • †Olinka Kukalová, 1958
  • ?†Paolekia Riek, 1976
  • †Paolia Smith, 1871
  • †Paoliola Handlirsch, 1919
  • †Permotermopsis Martynov, 1937
  • †Permula Handlirsch, 1919
  • †Protoblattina Meunier, 1909
  • †Pseudofouquea Handlirsch, 1906
  • †Silesiapteron Prokop et al., 2013
  • †Simplexpaolia Santos et al., 2023^[10]
  • †Sojanidelia Storozhenko, 1992
  • †Stenaropodites Martynov, 1928
  • †Sustaia Kukalová, 1958
  • †Sylvidelia Martynov, 1940
  • †Zdenekia Kukalová, 1958

References